Scent-Marking in Wolves

Foundational study on wolf scent marking behavior and degradation rates. The finding that scent marks degrade 30% per week (worthless after 3 weeks) forces constant boundary renewal - directly paralleling business need for continuous brand and relationship maintenance.

The 90% reduction in physical conflicts achieved through scent marking (120/year to 12/year) demonstrates that information is cheaper than combat, the core insight behind the Boundary Maintenance Audit framework.