Citation

Biological signals as handicaps

Alan Grafen

Journal of Theoretical Biology (1990)

TL;DR

Mathematical proof that handicap signaling is evolutionarily stable

Provides the mathematical formalization of Zahavi's handicap principle, demonstrating rigorously that costly signals can be evolutionarily stable honest communication systems. Essential for understanding the game-theoretic foundations of signaling.

Key Findings from Grafen (1990)

  • Mathematical proof that handicap signaling is evolutionarily stable
  • Cost-quality correlation is necessary for honesty
  • Equilibrium analysis of signaling systems

Related Mechanisms for Biological signals as handicaps